| 1980 - 1983: | B.Sc. with distinction, in food Technology from the Faculty of Agriculture, Hebrew University, Rehovot |
| 1985 - 1987: | M.Sc. in Microbiology with distinction
Thesis subject: Characterisation of relA mutations in Escherichia coli |
| 1988 - 1991: | Doctoral studies, the Institute of Microbiology, Hebrew University, Hadassah Medical School, Jerusalem
Supervisor: Prof. Gad Glaser Thesis subject: Studies of a global control system in prokaryotes mediated by the alarmon - ppGpp. |
| 1992 - 1995: | Postdoctoral work at the Laboratory of Prof. Alan Fersht, MRC unit for protein function and design, Cambridge, UK.
Subject: Intra and intermolecular interactions of barstar, the intercellular inhibitor of barnase from Bacillus amyloliquefaciens |
| 1996 - 2002: | Senior Scientist, Department of Biological Chemistry, Weizmann Institute of Science |
| 2002 - 2011 : | Associate Professor, Department of Biological Chemistry, Weizmann Institute of Science |
| 2012 : | Professor, Department of Biological Chemistry, Weizmann Institute of Science |