What DNA features, beyond the short DBD-bound motifs, are recognized by TFs? And which are recognized by the TF IDRs? Based on our recent findings, we hypothesize that DNA conformation dynamics hold the key for genomic specificity, by exposing transient helical structures compatible with IDR chemistry. We investigate the DNA-IDR targeting rules by combining massive parallel binding assays inside cells, mutations designed to modulate DNA sequence of conformation, and biophysical binding assays in cell-free systems.